Why are some AH00035 exceptions logged in Apache logs?

Issue

The logs are showing errors like:

[Thu Nov 28 08:27:34.399641 2019] [core:error] [pid 111111:tid 111111111111111] (13)Permission denied: [client 10.10.10.10:11111] AH00035: access to /path.to/file denied (filesystem path ‘/path.to/file’) because search permissions are missing on a component of the path, referer: https://www.example.com/sample.js
[Thu Nov 28 10:40:35.574493 2019] [core:error] [pid 1111:tid 111111111111111] (13)Permission denied: [client 10.10.10.10:11111] AH00035: access to /path.to/file denied (filesystem path ‘/path.to/file’) because search permissions are missing on a component of the path, referer: https://www.example.com/sample